Tait will start only his third Test
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England coach Andy Robinson insists Mathew Tait's future lies at outside centre - despite switching him to the wing against Australia on Saturday.
Tait replaces the dropped Tom Voyce on the left wing, with Newcastle team-mate Jamie Noon starting at outside centre.
"Mathew's future is as a 13," said Robinson, who made six changes in all.
"In the position we are in, and the players we had to call from for this game, I think it is better to involve him starting on the wing."
With James Simpson-Daniel, who was due to start the first Test before suffering a haematoma in his leg, out of the reckoning, Robinson had few other back-three options apart from full-back Mark van Gisbergen.
"I don't think Voyce gave us the penetration we needed (in the first Test)," Robinson explained. "I had to weigh up the choice between Voyce or Tom Varndell and, for me, I thought Varndell offered more.
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Chris Jones was quite an angry man at not being selected for the first Test
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"Yes, he (Varndell) made a couple of errors, but I certainly saw his cutting edge as important for this game.
"As a back line, we didn't have a lot of penetration in the second half, and I think Jamie Noon just gives that better balance for the penetration we are going to need."
Robinson also sprung by a surprise by dropping Olly Barkley at fly-half in favour of Leicester's Andy Goode, renown as more of a kicking number 10.
"I thought Andy did well when he came on (last Sunday)," Robinson explained. "We needed to attack the line a bit more with our 10, plus the range of our kicking skills."
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SCRUM V VIEW
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Robinson has also revamped his pack, with George Chuter replacing Lee Mears at hooker, Chris Jones and Ben Kay forming a new second row combination and Joe Worsley coming into the back row.
"A couple of changes were pre-planned, and a couple were because of the way we played," the coach added.
"I was looking at changes in the second row, with Ben Kay coming in. And Chris Jones was quite an angry man at not being selected for the first Test, but I thought his response was first-class.
"Chris was asked to get a tighter game, and I think he has worked hard at doing that. He has put on more weight, and he really punches his weight now.
"International rugby is a ruthless place, and we have got to be able to dominate opposition sides physically."