The four millionth Land Rover has rolled off the firm's production line in the West Midlands.
Land Rovers have been made at the plant since 1948
The iconic vehicles have been made at the firm's Solihull plant since 1948.
The latest Discovery 3 was collected by actress Joanna Lumley, who is a patron of the Born Free Foundation, a conservation charity.
The vehicle is being donated to the charity which will use it as a "rapid response rescue" vehicle for use in the UK and Europe.
It is estimated that 80% of all Land Rover Defenders ever produced are still in use, mainly on farms.
Land Rover managing director Phil Popham said: "We are delighted to hand this car over to Born Free.
"The production of the four millionth Land Rover is a major milestone and it is only fitting that it goes to such a worthy cause."
Born Free chief executive Will Travers said: "This important vehicle will enable us to carry out important frontline conservation and animal welfare missions at will.
"Without Land Rover's support we simply would not be able to reach some of the off-road remote destinations or tackle some of the rescues that are a vital part of our work."