Costs have risen even faster than income
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Higher education in the UK had an income of £18bn in the last academic year, latest statistics show.
This was 6.5% higher than the 2003-04 total of £16.9bn, according to the Higher Education Statistics Agency.
The biggest single source of income was grants from funding councils. But the biggest increase, almost 24%, came in the form of investments and donations.
Spending rose by 6.7% to £17.8bn, so the surplus within the sector overall fell from £237m to £213m.
The endowment and investment increase in Scotland was 11% compared with 53% for Northern Ireland, 32% for Welsh universities and nearly 25% in England.
A spokesman for the vice-chancellors' group, Universities UK, said the increase reflected in part the extra fundraising initiatives of recent years.
"Although fundraising is seen as a long term activity by higher education institutions and is only a small part of the overall income, it's clear that the higher priority given to it is beginning to pay off."
This year 27 had received matched funding from the government to increase their income from private donations.
Pay dispute
Staff costs rose from £9.7bn to £10.4bn.
Academic staff in two unions are engaged in an assessment and marking boycott in pursuit of higher pay.
The ongoing action follows a one-day strike in March.
A second round of talks, arranged through the conciliation service Acas, are set to take place on Tuesday.
The employers have offered a rise of 3% from August this year and an extra 3% from August 2007.
A two-day pay strike in further education colleges, planned for next month, has been suspended following a meeting between the union Natfhe and the employers, the Association of Colleges.
Natfhe spokesman Barry Lovejoy said: "We very much hope that when we next meet to discuss pay, lecturers will get a decent offer that means industrial action in colleges can be avoided this year."