A silverback gorilla from the jungles of Rwanda has died aged 35.
Titus found fame after he was filmed climbing on Sir David Attenborough's back in the 1979 BBC documentary Life on Earth.
Sir David said he was saddened by the news as 35 is a relatively young age for a gorilla to die. He added that the experience of meeting Titus had been a memorable moment which he would never be allowed to forget.
Titus was named by the American zoologist Dian Fossey, and he "starred" in a film about her - Gorillas in the Mist. He also featured in the BBC series The Gorilla King.